G.C. Williams, California Academy of Sciences (Last revised: 24 September 2019)
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[See: Williams, G.C. and P. Alderslade. 2011. Three new species of pennatulacean octocorals with the ability to attach to rocky substrata (Cnidaria: Anthozoa: Pennatulacea). Zootaxa 3001: 33-48]